📄 第六章.htm
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<h1 align=center style='text-align:center'><span style='font-family:隶书;
color:#EFCE8F'>第六章</span><span lang=EN-US style='font-size:36.0pt;mso-ascii-font-family:
隶书;mso-fareast-font-family:隶书;color:#EFCE8F'> </span><span
style='font-family:隶书;color:#EFCE8F'>系统发生分析</span><span lang=EN-US><o:p></o:p></span></h1>
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<p><span lang=EN-US><SELECT NAME="str_sel"
onchange="javascript:window.location=(this.options[this.selectedIndex].value);">
<OPTION SELECTED>========= 选择章节 ==========
<OPTION VALUE="6.1.htm">6.1 分子系统发生与系统发生树
<OPTION VALUE="6.2.htm">6.2 基于距离的系统发生树构建方法
<OPTION VALUE="6.3.htm">6.3 基于特征的系统发生树构建方法
<OPTION VALUE="6.4.htm">6.4 最大似然法
<OPTION VALUE="6.5.htm">6.5 系统发生树的可靠性
<OPTION VALUE="6.6.htm">6.6 全基因组系统发生分析
<OPTION VALUE="6.question.htm">问题与练习
<OPTION VALUE="6.referance.htm">参考文献
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<p style='line-height:150%'><span lang=EN-US style='font-size:10.5pt;
font-family:"Times New Roman";mso-font-kerning:1.0pt'> </span><span
style='font-size:10.5pt;mso-ascii-font-family:"Times New Roman";mso-hansi-font-family:
"Times New Roman";mso-font-kerning:1.0pt'>系统发生(或种系发生、系统发育,</span><span
lang=EN-US style='font-size:10.5pt;font-family:"Times New Roman";mso-font-kerning:
1.0pt'>phylogeny</span><span style='font-size:10.5pt;mso-ascii-font-family:
"Times New Roman";mso-hansi-font-family:"Times New Roman";mso-font-kerning:
1.0pt'>)是指生物形成或进化的历史。系统发生学</span><span lang=EN-US style='font-size:10.5pt;
font-family:"Times New Roman";mso-font-kerning:1.0pt'>(phylogenetics)</span><span
style='font-size:10.5pt;mso-ascii-font-family:"Times New Roman";mso-hansi-font-family:
"Times New Roman";mso-font-kerning:1.0pt'>研究物种之间的进化关系,其基本思想是比较物种的特征,并认为特征相似的物种在遗传学上接近。系统发生研究的结果往往以系统发生树(</span><span
lang=EN-US style='font-size:10.5pt;font-family:"Times New Roman";mso-font-kerning:
1.0pt'>phylogenetic tree</span><span style='font-size:10.5pt;mso-ascii-font-family:
"Times New Roman";mso-hansi-font-family:"Times New Roman";mso-font-kerning:
1.0pt'>)表示,用它描述物种之间的进化关系。通过对生物学数据的建模提取特征,进而比较这些特征,研究生物形成或进化的历史。在分子水平上进行系统发生分析具有许多优势,所得到的结果更加科学、可靠。分子系统发生分析主要分成三个步骤:分子序列或特征数据的分析、系统发生树的构造以及结果的检验。</span><span
lang=EN-US><o:p></o:p></span></p>
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<h2 align=center style='text-align:center'><!--mstheme--><span lang=EN-US
style='font-family:隶书;color:#EFCE8F'>6.1 分子系统发生与系统发生树 </span><span lang=EN-US><o:p></o:p></span></h2>
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<h3><!--mstheme--><span lang=EN-US style='font-size:12.0pt;color:#EFCE8F'>6.1.1
分子系统发生分析 </span><span lang=EN-US><o:p></o:p></span></h3>
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<p class=MsoNormal style='mso-margin-top-alt:auto;mso-margin-bottom-alt:auto;
text-indent:21.25pt;line-height:150%'><span style='font-size:10.0pt;font-family:
宋体;mso-ascii-font-family:"Times New Roman";mso-hansi-font-family:"Times New Roman"'>系统发生学是进化生物学的一个重要研究领域,系统发生分析早在达尔文时代就已经开始。从那时起,科学家们就开始寻找物种的源头,分析物种之间的进化关系,给各个物种分门别类。经典系统发生学研究所涉及的特征主要是生物表型</span><span
lang=EN-US style='font-size:10.0pt'>(phenotype)</span><span style='font-size:
10.0pt;font-family:宋体;mso-ascii-font-family:"Times New Roman";mso-hansi-font-family:
"Times New Roman"'>特征,所谓的表型特征主要指形态学的(结构的)特征,如生物体的大小、颜色、触角个数,也包括某些生理的、生化的以及行为习性的特征。通过表型比较来推断生物体的基因型</span><span
lang=EN-US style='font-size:10.0pt'>(genotype)</span><span style='font-size:
10.0pt;font-family:宋体;mso-ascii-font-family:"Times New Roman";mso-hansi-font-family:
"Times New Roman"'>,研究物种之间的进化关系。但是,利用表型特征是有局限性的。有时候关系很远的物种也能进化出相似的表型,这是由称为趋同进化</span><span
lang=EN-US style='font-size:10.0pt'>(convergent evolution)</span><span
style='font-size:10.0pt;font-family:宋体;mso-ascii-font-family:"Times New Roman";
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